Penholoway Swamp WMA consists of 4,269 acres owned and managed by the Georgia Department of Natural Resources (GA DNR) Nongame Section for wildlife, hunting, and other recreational activities. This WMA contains several stands which have been planted with longleaf pine (Pinus palustris) and is managed with prescribed fire and mechanical thinning of longleaf pine stands as needed.  The restoration of longleaf pine at this WMA is beneficial for a number of species such as gopher tortoises (Gopherus polyphemus). Gopher tortoises prefer deep sandy soils in which to dig their burrows. They also forage upon grasses, forbs, and other herbaceous groundcover that grow throughout the open understory that is iconic of historic longleaf pine habitat which used to dominate the southeastern United States.